This seems to be only an issue with the Exchange account. In case it matters, I'm running on an Early 2011 MBP with 8Gb RAM. I have also noticed that the unread count on the Inbox does not behave consistently: sometimes it is decremented even though the message still shows as Unread, sometimes the number does not change. For example, in this screenshot, I have clicked on all of these messages, and some have updated and some have not: This no longer happens reliably: some messages are marked as read, others are not. Before the Yosemite upgrade, whenever I selected the message summary in the left hand pane, it was instantly marked as "read". Using Yosemite (OSX 10.10) and the included Mail application (which describes itself as "Version 8.0 (1990.1)" to read mail held on a company Exchange server, I am finding that the unread marks for messages are not being reliably updated.
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